word-forming element, especially in scientific compounds, meaning "life, life and," or "biology, biology and," or "biological, of or pertaining to living organisms or their constituents," from Greek bios "one's life, course or way of living, lifetime" (as opposed to zoe "animal life, organic life"), from PIE root *gwei- "to live."
The correct usage is that in biography, but since c. 1800 in modern science it has been extended to mean "organic life," as zoo-, the better choice, is restricted in modern use to animal, as opposed to plant, life. Both are from the same PIE root. Compare biology.
word-forming element meaning "mind, mental; spirit, unconscious," from Greek combining form of psykhē "the soul, mind, spirit; life, one's life, the invisible animating principle or entity which occupies and directs the physical body; understanding, the mind (as the seat of thought), faculty of reason" (see psyche). It also was used to form compounds in Greek, such as psychapates "soul-beguiling" (with apate "deceit").
word-forming element meaning "animal, living being," from Greek zoion "an animal," literally "a living being," from PIE root *gwei- "to live" (source also of Greek bios "life").
word-forming element meaning "on the near side of, on this side," from Latin preposition cis "on this side" (in reference to place or time), related to citra (adv.) "on this side," from PIE *ki-s, suffixed form of root *ko-, the stem of demonstrative pronoun meaning "this." Opposed to trans- or ultra-. Originally only of place, sometimes 19c. of time; 21c. of life situations (such as cis-gender, which is attested by 2011).
element meaning "into, in, on, upon" (also im-, il-, ir- by assimilation of -n- with following consonant), from Latin in- "in," from PIE root *en "in."
In Old French (and hence in Middle English) this often became en-, which in English had a strong tendency to revert to Latin in-, but not always, which accounts for pairs such as enquire/inquire. There was a native form, which in West Saxon usually appeared as on- (as in Old English onliehtan "to enlighten"), and some of those verbs survived into Middle English (such as inwrite "to inscribe"), but all now seem to be extinct.
Not related to in- (1) "not," which also was a common prefix in Latin, causing confusion: to the Romans impressus could mean "pressed" or "unpressed;" inaudire meant "to hear," but inauditus meant "unheard of;" in Late Latin investigabilis could mean "that may be searched into" or "that cannot be searched into." Latin invocatus was "uncalled, uninvited," but invocare was "to call, appeal to."
The trouble has continued in English; the hesitation over what is meant by inflammable being a commonly cited example. Implume (1610s) meant "to feather," but implumed (c. 1600) meant "unfeathered." Impliable can mean "capable of being implied" (1865) or "inflexible" (1734). Impartible in 17c. could mean "incapable of being divided" or "capable of being imparted." Impassionate can be "free from passion" or it can mean "strongly stirred by passion." Inanimate (adj.) is "lifeless," but Donne uses inanimate (v.) to mean "infuse with life or vigor." Irruption is "a breaking in," but irruptible is "unbreakable."
In addition to improve "use to one's profit," Middle English also had a verb improve meaning "to disprove" (15c.). To inculpate is "to accuse," but inculpable means "not culpable, free from blame." Infestive has meant "troublesome, annoying" (1560s, from infest) and "not festive" (1620s). In Middle English inflexible could mean "incapable of being bent" or "capable of being swayed or moved." In 17c., informed could mean "current in information," formed, animated," or "unformed, formless" ("This was an awkward use" [OED]). Inhabited has meant "dwelt in" (1560s) and "uninhabited" (1610s); inhabitable likewise has been used on opposite senses, a confusion that goes back to Late Latin.
