Etymology
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*skoto- 

Proto-Indo-European root meaning "dark, shade." 

It forms all or part of: nightshade; scotoma; shade; shadow; shady.

It is the hypothetical source of/evidence for its existence is provided by: Greek skotos "darkness, gloom;" Albanian kot "darkness;" Old Irish scath, Old Welsh scod, Breton squeut "darkness," Gaelic sgath "shade, shadow, shelter;" Old English scead "partial darkness," sceadu "shade, shadow, darkness," Dutch schaduw, German Schatten, Gothic skadus "shadow." 

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*nebh- 
Proto-Indo-European root meaning "cloud."

It forms all or part of: nebula; nebular; nebulosity; nebulous; Neptune; Nibelungenlied; Niflheim; nimbus.

It is the hypothetical source of/evidence for its existence is provided by: Sanskrit nabhas- "vapor, cloud, mists, fog, sky;" Greek nephele, nephos "cloud;" Latin nebula "mist, vapor, fog, smoke, exhalation;" German Nebel "fog;" Old English nifol "dark, gloomy;" Welsh niwl "cloud, fog;" Slavic nebo.
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*bher- (2)

Proto-Indo-European root meaning "bright; brown" (the sense connection might involve polished wooden objects).

It forms all or part of: Barnard; bear (n.) "large carnivorous or omnivorous mammal of the family Ursidae;" beaver (n.1) "large amphibious quadruped rodent of the genus Castor;" berserk; brown; Bruin; brunet; brunette; burnish.

It is the hypothetical source of/evidence for its existence is provided by: Old English brun "dark, dusky;" Lithuanian bėras "brown;" Greek phrynos "toad," literally "the brown animal."

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*mori- 

Proto-Indo-European root meaning "body of water."

It forms all or part of: aquamarine; Armorica; beche-de-mer; cormorant; mare (n.2) "broad, dark areas of the moon;" marina; marinate; marine; mariner; maritime; marsh; mere (n.1) "lake, pool;" Merlin; mermaid; merman; meerschaum; meerkat; morass; Muriel; rosemary; submarine; ultramarine; Weimar.

It is the hypothetical source of/evidence for its existence is provided by: Latin mare; Old Church Slavonic morje, Russian more, Lithuanian marės, Old Irish muir, Welsh mor "sea;" Old English mere "sea, ocean; lake, pool," German Meer "sea."

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*pel- (1)

Proto-Indo-European root meaning "pale."

It forms all or part of: appall; falcon; fallow (adj.) "pale yellow, brownish yellow;" Fauvist; Lloyd; pale (adj.); pallid; pallor; palomino; Peloponnesus; polio; poliomyelitis.

It is the hypothetical source of/evidence for its existence is provided by: Sanskrit palitah "gray," panduh "whitish, pale;" Greek pelios "livid, dark;" polios "gray" (of hair, wolves, waves); Latin pallere "to be pale," pallidus "pale, pallid, wan, colorless;" Old Church Slavonic plavu, Lithuanian palvas "sallow;" Welsh llwyd "gray;" Old English fealo, fealu "dull-colored, yellow, brown." It also forms the root of words for "pigeon" in Greek (peleia), Latin (palumbes), and Old Prussian (poalis).

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*(s)keu- 

Proto-Indo-European root meaning "to cover, conceal."

It forms all or part of: chiaroscuro; cunnilingus; custody; cutaneous; cuticle; -cyte; cyto-; hide (v.1) "to conceal;" hide (n.1) "skin of a large animal;" hoard; hose; huddle; hut; kishke; lederhosen; meerschaum; obscure; scum; skewbald; skim; sky.

It is the hypothetical source of/evidence for its existence is provided by: Sanskrit kostha "enclosing wall," skunati "covers;" Greek kytos "a hollow, vessel," keutho "to cover, to hide," skynia "eyebrows;" Latin cutis "skin," ob-scurus "dark;" Lithuanian kiautas "husk," kūtis "stall;" Armenian ciw "roof;" Russian kishka "gut," literally "sheath;" Old English hyd "a hide, a skin," hydan "to hide, conceal; Old Norse sky "cloud;" Old English sceo "cloud;" Middle High German hode "scrotum;" Old High German scura, German Scheuer "barn;" Welsh cuddio "to hide."

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