Etymology
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-head 

word-forming element meaning "state or condition of being," Middle English -hede, from a variant of Old English -had, the source of -hood. The only surviving words with it are maidenhead and godhead.

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-headed 

"having a head" (of a specified kind); see head (n.).

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encephalo- 

before vowels encephal-, word-forming element meaning "brain, of the brain," from combining form of medical Latin encephalon, from Greek enkephalos "the brain," literally "within the head," from en "in" (see en- (2)) + kephalē "head" (see cephalo-).

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lepido- 

before vowels lepid-, word-forming element used since late 18c. in science with a sense of "scale" (of a fish, etc.), combining form of Greek lepis (genitive lepidos) "scale of a fish" (related to lepein "to peel;" see leper). As in lepidodendron (1819 in German), common fossil "club-moss tree" of the Carboniferous.

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-ful 

word-forming element attached to nouns (and in modern English to verb stems) and meaning "full of, having, characterized by," also "amount or volume contained" (handful, bellyful); from Old English -full, -ful, which is full (adj.) become a suffix by being coalesced with a preceding noun, but originally a separate word. Cognate with German -voll, Old Norse -fullr, Danish -fuld. Most English -ful adjectives at one time or another had both passive ("full of x") and active ("causing x; full of occasion for x") senses.

It is rare in Old English and Middle English, where full was much more commonly attached at the head of a word (for example Old English fulbrecan "to violate," fulslean "to kill outright," fulripod "mature;" Middle English had ful-comen "attain (a state), realize (a truth)," ful-lasting "durability," ful-thriven "complete, perfect," etc.).

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