before vowels spermat-, word-forming element meaning "seed, sperm," used from 1880s in scientific compounds, from Greek sperma (genitive spermatos "seed" of an animal or plant; see sperm).

word-forming element meaning "berry, seed," or something shaped like them, from Latinized form of Greek kokkos "a grain, a seed," especially "kermes-berry, gall of the kermes oak" (actually an insect), which yields scarlet dye, a word of unknown origin, perhaps from a non-Greek source.

before vowels gon-, word-forming element from Greek gonos "seed, that which engenders," from PIE *gon-o-, suffixed form of root *gene- "give birth, beget."

before vowels spor-, word-forming element meaning "spore," from Greek spora "a seed, a sowing," related to sporas "scattered, dispersed," sporos "sowing," from PIE *spor-, variant of root *sper- "to spread, sow" (see sparse).

before vowels angi-, word-forming element meaning "vessel of the body," now often "covered or enclosed by a seed or blood vessel," from Latinized form of Greek angeion "case, capsule, vessel of the body," diminutive of angos "vessel, jar, vat, vase," which is of unknown origin. Beekes says "Possibly a Mediterranean loanword ..., as kitchen utensils are often borrowed."

word-forming element in scientific compounds meaning "cartilage," from Latinized form of Greek khondros "cartilage" (of the breastbone), also "grain, grain of salt, seed, barley-grain," of uncertain origin. This is sometimes said to be from the PIE root meaning "to grind" which is the source of English grind (v.), but there are serious phonological objections and the word might be non-Indo-European [Beekes, "Etymological Dictionary of Greek"]. The body material so called for its gristly nature.