c. 1300, from Old French veine "vein, artery, pulse" (12c.), from Latin vena "a blood vessel," also "a water course, a vein of metal, a person's natural ability or interest," of unknown origin. The mining sense is attested in English from late 14c. (Greek phleps "vein" had the same secondary sense). Figurative sense of "strain or intermixture" (of some quality) is recorded from 1560s; that of "a humor or mood, natural tendency" is first recorded 1570s.
word-forming element in medicine meaning "a vein or veins," from Greek phlebo-, combining form of phleps "vein," a word of uncertain origin.
"arrangement of veins," 1640s, of plant structures, noun of state from Latin vena "vein" (see vein). Related: Venational.
plural of varix "dilated vein" (c. 1400), from Latin varix "a varicose vein," which de Vaan derives from varus "bent outward, bow-legged," which is of uncertain origin (see vary).