word-forming element meaning "medical treatment," from French -iatrie, from Greek iatreia "healing, medical treatment" (see -iatric).

before vowels, hipp-, word-forming element meaning "horse," from Greek hippo-, from hippos "horse," from PIE root *ekwo- "horse."

word-forming element meaning "wrist," from combining form of Latin carpus, from Greek karpos "wrist" (see carpus).

word-forming element meaning "nature, natural, physical," from Greek physios "nature" (from PIE root *bheue- "to be, exist, grow").

word-forming element meaning "angle, corner," from Greek gōnia "corner, angle," from PIE root *genu- (1) "knee; angle."

word-forming element meaning "fruit," from Latinized form of Greek karpos "fruit," from PIE root *kerp- "to gather, pluck, harvest."

word-forming element meaning "drug, medicine," also "poison," from Latinized form of Greek pharmakon "drug, poison" (see pharmacy).

word-forming element making ordinal numbers (fourth, tenth, etc.), Old English -ða, from Proto-Germanic *-tha- (cognates: Gothic -da, -ta, Old High German -do, -to, Old Norse -di, -ti), from PIE *-to-, also *-eto-, *-oto-, suffix forming adjectives "marking the accomplishment of the notion of the base" [Watkins].

Cognate with Sanskrit thah, Greek -tos, Latin -tus; Sanskrit ta-, Lithuanian and Old Church Slavonic to, Greek to "the," Latin talis "such;" Greek tēlikos "so old, of such an age," Old Church Slavonic toli "so, to such a degree," toliku "so much," Russian toliko "only;" also see -ed.

word-forming element meaning "that which turns," from Greek tropos "a turn, direction, course, way," from PIE root *trep- "to turn."

before vowels spondyl-, combining form meaning "vertebrae," from Greek spondylos "a vertebra," in plural "the backbone," variant of sphondylos, of uncertain origin.